特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)(long)是發現于(yu)亞洲(zhou)地區(qu)的(de)(de)大型(xing)暴(bao)龍(long)(long)(long)(long)(long)(long)(long)類,在(zai)外形上與北(bei)美洲(zhou)的(de)(de)霸(ba)王龍(long)(long)(long)(long)(long)(long)(long)非(fei)常(chang)相似,是當時(shi)生態系(xi)統中的(de)(de)頂級捕食者。特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)(long)的(de)(de)體型(xing)比霸(ba)王龍(long)(long)(long)(long)(long)(long)(long)和諸城暴(bao)龍(long)(long)(long)(long)(long)(long)(long)略小,但明(ming)顯(xian)大于(yu)其它(ta)的(de)(de)暴(bao)龍(long)(long)(long)(long)(long)(long)(long)類,其頭骨(gu)的(de)(de)最大長度可(ke)達1.3米。特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)(long)的(de)(de)化(hua)石主要發現于(yu)蒙古(gu),但是也(ye)有一些證據(ju)表明(ming)這種巨(ju)型(xing)掠食者在(zai)中國的(de)(de)內蒙古(gu)以及新疆等(deng)地區(qu)都有分布(bu)。如同它(ta)在(zai)北(bei)美洲(zhou)的(de)(de)近(jin)親一樣,特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)(long)的(de)(de)標本量也(ye)非(fei)常(chang)巨(ju)大。截止(zhi)到目前為止(zhi),古(gu)生物學家(jia)至少已(yi)經(jing)找(zhao)到了約30個(ge)特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)(long)個(ge)體的(de)(de)標本,這其中不乏保存精美的(de)(de)頭骨(gu)化(hua)石。在(zai)亞洲(zhou)恐龍(long)(long)(long)(long)(long)(long)(long)古(gu)生物學發展的(de)(de)早期(qi)(qi)階段,特(te)(te)暴(bao)龍(long)(long)(long)(long)(long)(long)(long)不同發育(yu)階段的(de)(de)個(ge)體曾經(jing)被(bei)當成過不同的(de)(de)物種,不過在(zai)近(jin)期(qi)(qi)的(de)(de)研(yan)究中這些錯(cuo)誤(wu)都被(bei)一一糾正。
特(te)(te)暴(bao)龍(long)的(de)屬(shu)(shu)名意為(wei)“可怕的(de)蜥蜴”,種(zhong)名意為(wei)“勇(yong)士”。特(te)(te)暴(bao)龍(long)的(de)正型標(biao)本(ben)最初并(bing)沒(mei)有被(bei)歸(gui)入(ru)(ru)(ru)到特(te)(te)暴(bao)龍(long)屬(shu)(shu),而是被(bei)歸(gui)入(ru)(ru)(ru)到暴(bao)龍(long)屬(shu)(shu),并(bing)命名為(wei)“Tyrannosaurus bataar”,另外(wai)三個早期(qi)的(de)歸(gui)入(ru)(ru)(ru)標(biao)本(ben)中有兩個被(bei)歸(gui)入(ru)(ru)(ru)了蛇發女怪龍(long)屬(shu)(shu)。直到1965年蘇(su)聯古生物學家阿納(na)托利·康斯(si)坦丁諾維奇·羅特(te)(te)杰斯(si)特(te)(te)文斯(si)基才發現馬列(lie)夫命名的(de)這4件標(biao)本(ben)是同(tong)一物種(zhong)不同(tong)發育階段的(de)個體,隨后它們才被(bei)并(bing)入(ru)(ru)(ru)勇(yong)士特(te)(te)暴(bao)龍(long)。
正型標本:PIN 551-1保存了一(yi)部(bu)分(fen)頭骨和頸(jing)椎(zhui)。最(zui)早(zao)被歸入暴龍屬,后歸入特暴龍屬。
此(ci)(ci)外蘭平暴龍(long)(Tyrannosaurus lanpingensis),吐(tu)魯番暴龍(long)(Tyrannosaurus turpanensis),Albertosaurus periculosus目前都被認為是屬于勇士特(te)暴龍(long)的(de)(de)(de)不同個(ge)體(ti)發育(yu)階段。除(chu)此(ci)(ci)之(zhi)外,在蒙古(gu)發現的(de)(de)(de)分支龍(long)曾經也被懷疑過(guo)是特(te)暴龍(long)的(de)(de)(de)幼年(nian)個(ge)體(ti),不過(guo)近年(nian)來由于虔州龍(long)的(de)(de)(de)發現,分支龍(long)類的(de)(de)(de)有效性已經被學術界所接(jie)受。
特暴(bao)龍(long)的(de)(de)(de)(de)(de)(de)(de)頭(tou)骨(gu)側視與霸王(wang)(wang)龍(long)非(fei)常相(xiang)似。兩者都具有(you)背腹向(xiang)加深的(de)(de)(de)(de)(de)(de)(de)頭(tou)骨(gu),且(qie)具有(you)粗(cu)壯的(de)(de)(de)(de)(de)(de)(de)下頜(he)(he)。從(cong)背視可以看出兩種恐龍(long)的(de)(de)(de)(de)(de)(de)(de)頭(tou)骨(gu)整體輪廓的(de)(de)(de)(de)(de)(de)(de)差(cha)異,霸王(wang)(wang)龍(long)的(de)(de)(de)(de)(de)(de)(de)后端向(xiang)外側極度的(de)(de)(de)(de)(de)(de)(de)擴(kuo)展,使得(de)雙眼視覺范圍具有(you)較大的(de)(de)(de)(de)(de)(de)(de)重疊。雖然特暴(bao)龍(long)的(de)(de)(de)(de)(de)(de)(de)頭(tou)骨(gu)后端也有(you)一(yi)定程(cheng)度的(de)(de)(de)(de)(de)(de)(de)向(xiang)外擴(kuo)展,但(dan)是與霸王(wang)(wang)龍(long)有(you)較大的(de)(de)(de)(de)(de)(de)(de)差(cha)距(ju)。同時特暴(bao)龍(long)的(de)(de)(de)(de)(de)(de)(de)吻端明顯(xian)比霸王(wang)(wang)龍(long)更(geng)窄。霸王(wang)(wang)龍(long)的(de)(de)(de)(de)(de)(de)(de)前(qian)頜(he)(he)骨(gu)和上頜(he)(he)骨(gu)相(xiang)接的(de)(de)(de)(de)(de)(de)(de)骨(gu)縫上有(you)一(yi)個(ge)明顯(xian)的(de)(de)(de)(de)(de)(de)(de)開(kai)孔暴(bao)露在頭(tou)骨(gu)的(de)(de)(de)(de)(de)(de)(de)外側面,而這(zhe)一(yi)開(kai)孔在特暴(bao)龍(long)頭(tou)骨(gu)的(de)(de)(de)(de)(de)(de)(de)外側面幾乎沒有(you)。
特暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)上(shang)(shang)頜(he)骨(gu)升突粗狀發達(da),霸(ba)(ba)王(wang)龍(long)(long)(long)的(de)(de)(de)(de)較為纖細。兩種恐龍(long)(long)(long)的(de)(de)(de)(de)鼻骨(gu)腹(fu)外(wai)(wai)側緣都具有與(yu)上(shang)(shang)頜(he)骨(gu)相(xiang)關節(jie)的(de)(de)(de)(de)溝,并幾(ji)乎前(qian)后向延伸至鼻骨(gu)的(de)(de)(de)(de)整個長度。但特暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)關節(jie)面具有很(hen)多脊,使鼻骨(gu)與(yu)上(shang)(shang)頜(he)骨(gu)的(de)(de)(de)(de)關節(jie)更(geng)緊(jin)密(mi),而(er)霸(ba)(ba)王(wang)龍(long)(long)(long)的(de)(de)(de)(de)鼻骨(gu)上(shang)(shang)頜(he)骨(gu)關節(jie)更(geng)簡單(dan)。相(xiang)反的(de)(de)(de)(de)是,霸(ba)(ba)王(wang)龍(long)(long)(long)的(de)(de)(de)(de)鼻骨(gu)與(yu)淚骨(gu)的(de)(de)(de)(de)關節(jie)十分復雜(za)且(qie)非(fei)常緊(jin)密(mi),而(er)特暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)鼻骨(gu)與(yu)淚骨(gu)的(de)(de)(de)(de)關節(jie)則較為簡單(dan)。此外(wai)(wai),霸(ba)(ba)王(wang)龍(long)(long)(long)的(de)(de)(de)(de)鼻骨(gu)比特暴(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)更(geng)加寬大。
霸(ba)王龍的(de)(de)(de)淚(lei)(lei)骨背(bei)緣較特暴龍有更(geng)明顯的(de)(de)(de)膨大。特暴龍的(de)(de)(de)淚(lei)(lei)骨降支(zhi)較為(wei)中空(kong),而霸(ba)王龍的(de)(de)(de)淚(lei)(lei)骨降支(zhi)幾乎(hu)是實(shi)心(xin)的(de)(de)(de)。特暴龍的(de)(de)(de)淚(lei)(lei)骨前突(tu)與霸(ba)王龍的(de)(de)(de)相(xiang)比更(geng)為(wei)細長,且前突(tu)關(guan)(guan)節面(mian)沒(mei)有分支(zhi)。霸(ba)王龍的(de)(de)(de)淚(lei)(lei)骨前突(tu)關(guan)(guan)節面(mian)具(ju)(ju)有明顯的(de)(de)(de)分叉(cha)。淚(lei)(lei)骨前突(tu)的(de)(de)(de)內側面(mian)中段具(ju)(ju)有一(yi)個容納血(xue)管的(de)(de)(de)開孔,這(zhe)一(yi)開孔在(zai)霸(ba)王龍的(de)(de)(de)淚(lei)(lei)骨上的(de)(de)(de)位置更(geng)靠前。
鱗骨(gu)(gu)(gu)(gu)的(de)(de)(de)(de)(de)降支與方(fang)顴骨(gu)(gu)(gu)(gu)的(de)(de)(de)(de)(de)上支具(ju)有很大的(de)(de)(de)(de)(de)接觸面(mian)(mian)(mian)(mian)積,與艾(ai)伯塔(ta)龍(long)(long)(long)(long)和(he)蛇(she)發(fa)女怪龍(long)(long)(long)(long)不同,而(er)與霸(ba)(ba)王龍(long)(long)(long)(long)非(fei)常相似。在(zai)腹側(ce)(ce)視,特暴(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)顴骨(gu)(gu)(gu)(gu)不如(ru)霸(ba)(ba)王龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)那樣(yang)很明(ming)(ming)顯的(de)(de)(de)(de)(de)外(wai)翻(fan)。特暴(bao)龍(long)(long)(long)(long)和(he)霸(ba)(ba)王龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)顴骨(gu)(gu)(gu)(gu)升突(tu)外(wai)側(ce)(ce)面(mian)(mian)(mian)(mian)都(dou)具(ju)有一(yi)(yi)(yi)(yi)不甚明(ming)(ming)顯的(de)(de)(de)(de)(de)凹(ao)陷,但是霸(ba)(ba)王龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)這一(yi)(yi)(yi)(yi)凹(ao)陷明(ming)(ming)顯小于(yu)(yu)特暴(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)。兩者的(de)(de)(de)(de)(de)顴骨(gu)(gu)(gu)(gu)后突(tu)都(dou)具(ju)有二分支的(de)(de)(de)(de)(de)結(jie)構,區(qu)別在(zai)于(yu)(yu)特暴(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)顴骨(gu)(gu)(gu)(gu)后突(tu)腹側(ce)(ce)支的(de)(de)(de)(de)(de)厚(hou)度不均(jun)(jun)勻(yun),其腹緣較厚(hou),向(xiang)背緣發(fa)展的(de)(de)(de)(de)(de)過程中逐漸變薄,而(er)霸(ba)(ba)王龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)顴骨(gu)(gu)(gu)(gu)后突(tu)腹側(ce)(ce)支的(de)(de)(de)(de)(de)厚(hou)度更(geng)為均(jun)(jun)一(yi)(yi)(yi)(yi),且(qie)(qie)外(wai)側(ce)(ce)面(mian)(mian)(mian)(mian)更(geng)為明(ming)(ming)顯的(de)(de)(de)(de)(de)隆起(qi)。特暴(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)方(fang)顴骨(gu)(gu)(gu)(gu)柄明(ming)(ming)顯比(bi)(bi)霸(ba)(ba)王龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)更(geng)加纖弱(ruo)(ruo)。霸(ba)(ba)王龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)方(fang)顴骨(gu)(gu)(gu)(gu)前突(tu)比(bi)(bi)特暴(bao)龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)更(geng)為寬大,能夠覆蓋住顴骨(gu)(gu)(gu)(gu)后突(tu)的(de)(de)(de)(de)(de)大部分面(mian)(mian)(mian)(mian)積,且(qie)(qie)內側(ce)(ce)面(mian)(mian)(mian)(mian)具(ju)有一(yi)(yi)(yi)(yi)個很大的(de)(de)(de)(de)(de)凹(ao)陷。特暴(bao)龍(long)(long)(long)(long)顎骨(gu)(gu)(gu)(gu)上的(de)(de)(de)(de)(de)內鼻孔(kong)邊緣發(fa)育的(de)(de)(de)(de)(de)很明(ming)(ming)顯,而(er)這一(yi)(yi)(yi)(yi)特征在(zai)霸(ba)(ba)王龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)顎骨(gu)(gu)(gu)(gu)上相對較弱(ruo)(ruo)。特暴(bao)龍(long)(long)(long)(long)外(wai)翼骨(gu)(gu)(gu)(gu)上的(de)(de)(de)(de)(de)氣孔(kong)比(bi)(bi)霸(ba)(ba)王龍(long)(long)(long)(long)的(de)(de)(de)(de)(de)更(geng)小。
特(te)(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)犁骨(gu)(gu)(gu)前(qian)(qian)突(tu)向(xiang)前(qian)(qian)極(ji)度(du)延伸,與前(qian)(qian)頜骨(gu)(gu)(gu)的(de)顎(e)突(tu)相接,而(er)這一(yi)(yi)(yi)(yi)特(te)(te)(te)征在霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)中(zhong)不(bu)明顯。特(te)(te)(te)暴(bao)(bao)龍(long)(long)(long)和(he)霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)翼骨(gu)(gu)(gu)的(de)最大(da)區別在于(yu)(yu)顎(e)骨(gu)(gu)(gu)關節面(mian)的(de)前(qian)(qian)邊(bian)緣(yuan)。霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)的(de)翼骨(gu)(gu)(gu)額骨(gu)(gu)(gu)突(tu)的(de)前(qian)(qian)邊(bian)緣(yuan)是一(yi)(yi)(yi)(yi)條(tiao)斜(xie)線,而(er)特(te)(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)翼骨(gu)(gu)(gu)顎(e)骨(gu)(gu)(gu)突(tu)前(qian)(qian)邊(bian)緣(yuan)向(xiang)前(qian)(qian)突(tu)出(chu)。特(te)(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)上枕骨(gu)(gu)(gu)的(de)內外向(xiang)寬(kuan)度(du)小(xiao)于(yu)(yu)枕髁(ke),而(er)霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)的(de)上枕骨(gu)(gu)(gu)更(geng)寬(kuan)。從側(ce)面(mian)看,特(te)(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)齒(chi)骨(gu)(gu)(gu)吻(wen)端比霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)的(de)更(geng)加尖細一(yi)(yi)(yi)(yi)點。霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)的(de)夾板(ban)骨(gu)(gu)(gu)腹緣(yuan)相較于(yu)(yu)特(te)(te)(te)暴(bao)(bao)龍(long)(long)(long)有(you)更(geng)大(da)的(de)傾(qing)斜(xie)角(jiao)度(du),并(bing)且夾板(ban)骨(gu)(gu)(gu)的(de)前(qian)(qian)端位于(yu)(yu)更(geng)高(gao)的(de)位置(zhi)。特(te)(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)上隅骨(gu)(gu)(gu)孔較霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)的(de)要(yao)發達(da)。霸(ba)(ba)王(wang)(wang)(wang)龍(long)(long)(long)和(he)特(te)(te)(te)暴(bao)(bao)龍(long)(long)(long)的(de)成(cheng)體上頜骨(gu)(gu)(gu)齒(chi)和(he)齒(chi)骨(gu)(gu)(gu)齒(chi)的(de)數(shu)量不(bu)一(yi)(yi)(yi)(yi)樣。
歸(gui)入標本:PIN 551-2是一具接近完整的(de)骨架。最(zui)早(zao)被(bei)歸(gui)入到特(te)暴(bao)龍屬(shu)的(de)歸(gui)入種(zhong)(zhong)埃夫雷莫夫特(te)暴(bao)龍(Tarbosaurus efremovi)。現已(yi)歸(gui)入到勇士特(te)暴(bao)龍,且(qie)勇士特(te)暴(bao)龍是特(te)暴(bao)龍屬(shu)下的(de)唯一有效(xiao)種(zhong)(zhong)。
歸(gui)(gui)入標(biao)本:PIN 553-1保存了(le)頭骨(gu)(gu),背椎和尾椎,掌(zhang)骨(gu)(gu)和跖骨(gu)(gu)。最開始被歸(gui)(gui)入到蛇發女怪龍屬(shu)。現已歸(gui)(gui)入勇(yong)士(shi)特暴龍。
歸(gui)入(ru)標(biao)本:PIN 552-2保存了頭骨和頭后骨骼。最開始被歸(gui)入(ru)到蛇發女(nv)怪龍屬。現已歸(gui)入(ru)勇士特暴龍。
歸入標本(ben)(ben):MPC-D 107/7是一件幼年的標本(ben)(ben),關聯保(bao)存。除了缺少頸椎(zhui),前(qian)部(bu)背椎(zhui)和末端尾椎(zhui),其它骨(gu)頭基本(ben)(ben)都有保(bao)存。
歸入標本:ZPAL MgD-I/3幾乎完(wan)整(zheng)的(de)中等體型的(de)個體。保存了頭骨(gu),頸椎和(he)(he)背椎,10節(jie)近端(duan)尾椎,完(wan)整(zheng)的(de)腸骨(gu),恥(chi)骨(gu)和(he)(he)坐骨(gu),肋(lei)骨(gu)和(he)(he)腹膜肋(lei),完(wan)整(zheng)的(de)肩胛骨(gu),烏(wu)喙骨(gu),左側的(de)前肢,不完(wan)整(zheng)保存的(de)右(you)側后(hou)肢。
歸入標本:ZPAL Mgd-I/4部分保(bao)存的大型個體。保(bao)存了左側(ce)后(hou)肢(zhi),腸骨,13節椎體。
歸入標本:ZPAL Mgd-I/5不完整(zheng)保(bao)存(cun)的(de)大型骨架。保(bao)存(cun)了左(zuo)(zuo)側(ce)(ce)上頜骨,左(zuo)(zuo)側(ce)(ce)方(fang)骨,左(zuo)(zuo)側(ce)(ce)的(de)下(xia)頜和右(you)側(ce)(ce)的(de)下(xia)頜碎(sui)片,11節左(zuo)(zuo)側(ce)(ce)肋骨的(de)碎(sui)片,恥骨碎(sui)片,坐骨,腸骨碎(sui)片,左(zuo)(zuo)側(ce)(ce)后肢和右(you)側(ce)(ce)跖骨,和很多碎(sui)屑。
歸入標本:ZPAL Mgd-I/26不(bu)完整保存的(de)上頜骨(gu)和一部分保存下來的(de)牙齒。
歸入標本:ZPAL Mgd-I/29不完(wan)整(zheng)的(de)(de)大型(xing)骨(gu)(gu)架。保(bao)存(cun)了頭(tou)骨(gu)(gu)和(he)較(jiao)為完(wan)整(zheng)的(de)(de)齒列。頭(tou)后骨(gu)(gu)骼保(bao)存(cun)了6節(jie)頸椎,5節(jie)薦椎,22節(jie)尾椎,11節(jie)右側(ce)肋(lei)骨(gu)(gu),腸骨(gu)(gu),不完(wan)整(zheng)的(de)(de)恥骨(gu)(gu)和(he)近端坐(zuo)骨(gu)(gu),左(zuo)側(ce)肱骨(gu)(gu),尺(chi)骨(gu)(gu)和(he)橈骨(gu)(gu)的(de)(de)遠(yuan)端,第一手指,幾乎完(wan)整(zheng)的(de)(de)右側(ce)后肢。
歸入標本:ZPAL Mgd-I/31右(you)側上(shang)頜骨的(de)近端。
歸(gui)入標本:ZPAL Mgd-I/34頭骨右側的碎(sui)片。
歸入(ru)標(biao)本:ZPAL Mgd-I/38不完全保存的(de)(de)頭骨(gu),12節(jie)(jie)肋骨(gu),右側(ce)股骨(gu)的(de)(de)遠端(duan)(duan)。右側(ce)脛骨(gu)的(de)(de)遠端(duan)(duan),右側(ce)的(de)(de)3-4跖骨(gu),第4趾(zhi)的(de)(de)近端(duan)(duan)趾(zhi)節(jie)(jie)骨(gu)。
歸入標(biao)本:ZPAL Mgd-I/44頭(tou)骨的(de)右側。
歸入標本:ZPAL Mgd-I/45頭骨碎片。
歸入標(biao)本:ZPAL Mgd-I/46右側下頜碎(sui)(sui)片和其它7塊頭骨碎(sui)(sui)片,和兩節斷(duan)掉的背椎肋骨。
歸(gui)入標本:ZPAL Mgd-I/52左側下頜齒(chi)骨。
歸入標本:ZPAL Mgd-I/67右(you)側顴(quan)骨。
歸(gui)入標本(ben):ZPAL Mgd-I/93頭骨腦顱(lu)。
歸入標本:ZPAL Mgd-I/109大的帶有(you)圍巖的頭骨。
歸入(ru)標本:ZPAL Mgd-I/178頭骨(gu)的碎片,脊(ji)椎骨(gu)和股骨(gu)。
歸入(ru)標本(ben):GIN 100/61頭(tou)骨(gu)碎(sui)片和(he)頭(tou)后骨(gu)骼(ge)。
歸入標本:GIN 100/62頭骨碎片和頭后骨骼(ge)。
歸入標(biao)本:GIN 100/65頭骨的右半部分(fen)。
歸入標本:GIN 100/67頭骨的碎片,腦顱。
歸(gui)入標本:GIN 100/69枕區。
歸入標本:GIN 100/70頭(tou)骨碎片和椎體。
歸入標(biao)本(ben):GIN 107/2完(wan)整的骨架。
歸入(ru)標本:GIN 107/3頭骨。
歸入標本:PIN 551-3骨架,保存狀況未(wei)描述(shu)。最早被歸入到特(te)暴龍(long)(long)(long)屬的歸入種埃夫(fu)雷莫(mo)夫(fu)特(te)暴龍(long)(long)(long)。現已歸入到勇士特(te)暴龍(long)(long)(long)。
歸(gui)入(ru)(ru)(ru)標本:PIN 551-4骨(gu)架,保存狀況(kuang)未描述。最早(zao)被(bei)歸(gui)入(ru)(ru)(ru)到(dao)特(te)(te)(te)暴龍屬的歸(gui)入(ru)(ru)(ru)種埃夫雷(lei)莫(mo)夫特(te)(te)(te)暴龍。現已歸(gui)入(ru)(ru)(ru)到(dao)勇士特(te)(te)(te)暴龍。
歸(gui)入(ru)標本:PIN 551-91右側上頜骨。最早被歸(gui)入(ru)到(dao)特暴(bao)龍(long)(long)屬的歸(gui)入(ru)種埃夫雷(lei)莫夫特暴(bao)龍(long)(long)。現已歸(gui)入(ru)到(dao)勇士特暴(bao)龍(long)(long)。
歸(gui)入標本(ben):PIN 552-1目前僅有(you)復(fu)制品保存。最早被歸(gui)入到(dao)特暴龍(long)屬的歸(gui)入種埃夫(fu)雷莫夫(fu)特暴龍(long)。現已歸(gui)入到(dao)勇士(shi)特暴龍(long)。
歸(gui)(gui)(gui)(gui)入(ru)標本:PIN 553-2保存狀況未描述。早被歸(gui)(gui)(gui)(gui)入(ru)到特(te)暴(bao)龍屬的歸(gui)(gui)(gui)(gui)入(ru)種(zhong)埃夫(fu)雷(lei)莫夫(fu)特(te)暴(bao)龍。現已歸(gui)(gui)(gui)(gui)入(ru)到勇士特(te)暴(bao)龍。
歸入標(biao)本(ben):IVPP V 4878火焰山鄯善龍(long)(Shanshanosaurus huoyanshanensis)的正型標(biao)本(ben),發現于中國(guo)新疆,現在認為(wei)可能是特暴龍(long)的幼體。保存(cun)了上頜(he)骨(gu),下頜(he)大部(bu)分骨(gu)塊(kuai),多節(jie)頸椎,背椎和(he)薦(jian)椎,肩帶骨(gu)骼,肱骨(gu),恥(chi)骨(gu)遠端,股骨(gu)和(he)脛骨(gu)近端。
歸(gui)入標(biao)本:IVPP V 4733欒川暴龍(long)(Tyrannosaurus luanchuanensis)的正型標(biao)本,為5顆牙齒,現認為也可能屬于特暴龍(long)。
歸入標本:IVPP V 836為(wei)破(po)碎金剛(gang)口龍(Chingkankousaurus fragilis)的正型(xing)標本,為(wei)一節不完整的肩胛骨標本。布魯薩特等人認為(wei)很有(you)可(ke)能是特暴(bao)龍,但不太確定。
特(te)暴(bao)龍(long)(long)同霸(ba)王龍(long)(long)一(yi)樣,是暴(bao)龍(long)(long)超科中最進步(bu)的成員。大多(duo)數的系統發育分析都支持(chi)霸(ba)王龍(long)(long)和特(te)暴(bao)龍(long)(long)構成一(yi)個姐妹(mei)群,諸城暴(bao)龍(long)(long)作為兩者所構成單系的姐妹(mei)群。也有(you)一(yi)些(xie)分析認(ren)為特(te)暴(bao)龍(long)(long)與(yu)諸城暴(bao)龍(long)(long)的親緣關系更(geng)近。
如果不考慮標本量(liang)很少且質量(liang)很差的(de)諸城暴(bao)(bao)(bao)龍(long)(long)(long),特暴(bao)(bao)(bao)龍(long)(long)(long)的(de)體型基(ji)本穩坐暴(bao)(bao)(bao)龍(long)(long)(long)類的(de)第二把交(jiao)椅。最大的(de)特暴(bao)(bao)(bao)龍(long)(long)(long)頭(tou)骨長度(du)達到(dao)1.3米,這一大小顯著的(de)大于其(qi)它的(de)大型暴(bao)(bao)(bao)龍(long)(long)(long)類,并基(ji)本達到(dao)了(le)個別霸王龍(long)(long)(long)成(cheng)年標本的(de)頭(tou)骨長度(du)。目(mu)前最大的(de)特暴(bao)(bao)(bao)龍(long)(long)(long)標本體長是肯(ken)定能超過(guo)10米的(de)。特暴(bao)(bao)(bao)龍(long)(long)(long)的(de)體重一般估計(ji)在4噸(dun)左右。
通(tong)過(guo)仔細對比(bi)霸王龍(long)(long)(long)(long)(long)及其它(ta)北美(mei)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)(long)亞科成員的(de)頭骨(gu)(gu)(gu)各個骨(gu)(gu)(gu)塊(kuai)骨(gu)(gu)(gu)縫與(yu)特暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)差異(yi)可以發現霸王龍(long)(long)(long)(long)(long)與(yu)特暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)頭骨(gu)(gu)(gu)具有不太一(yi)樣(yang)的(de)生物力(li)(li)(li)(li)學結(jie)構(gou)。大型(xing)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)(long)類(lei)都具有毀(hui)滅(mie)性的(de)咬(yao)合力(li)(li)(li)(li),而它(ta)們(men)的(de)頭骨(gu)(gu)(gu)也(ye)(ye)必(bi)須在撕咬(yao)獵(lie)物的(de)同時承受同樣(yang)巨大的(de)反(fan)作(zuo)用力(li)(li)(li)(li)。北美(mei)洲(zhou)的(de)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)(long)亞科主要通(tong)過(guo)上頜(he)骨(gu)(gu)(gu)-鼻骨(gu)(gu)(gu)-淚(lei)骨(gu)(gu)(gu)-額(e)骨(gu)(gu)(gu)/前(qian)額(e)骨(gu)(gu)(gu)的(de)緊(jin)密關節(jie)(jie)構(gou)成來承受撕咬(yao)獵(lie)物過(guo)程中(zhong)(zhong)產生的(de)反(fan)作(zuo)用力(li)(li)(li)(li)。其中(zhong)(zhong),鼻骨(gu)(gu)(gu)與(yu)淚(lei)骨(gu)(gu)(gu)構(gou)成了(le)非常強的(de)關節(jie)(jie),且這一(yi)特征普遍存在于北美(mei)的(de)大型(xing)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)(long)亞科中(zhong)(zhong)。對于特暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)研(yan)究顯示,與(yu)北美(mei)洲(zhou)的(de)親戚們(men)相比(bi),特暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)淚(lei)骨(gu)(gu)(gu)前(qian)突(tu)向下偏(pian)轉的(de)更多,并且擁(yong)有比(bi)北美(mei)洲(zhou)暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)(long)類(lei)更為強壯的(de)上頜(he)骨(gu)(gu)(gu)升突(tu),使得這兩個塊(kuai)骨(gu)(gu)(gu)頭通(tong)過(guo)復雜的(de)關節(jie)(jie)面(mian)緊(jin)密關節(jie)(jie)在一(yi)起。同時特暴(bao)(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)前(qian)額(e)骨(gu)(gu)(gu)與(yu)淚(lei)骨(gu)(gu)(gu)也(ye)(ye)有非常緊(jin)密的(de)關節(jie)(jie)。非常強大的(de)上頜(he)骨(gu)(gu)(gu)-淚(lei)骨(gu)(gu)(gu)關節(jie)(jie)也(ye)(ye)見于異(yi)特龍(long)(long)(long)(long)(long)和中(zhong)(zhong)華(hua)盜龍(long)(long)(long)(long)(long)這樣(yang)的(de)大型(xing)異(yi)特龍(long)(long)(long)(long)(long)類(lei)中(zhong)(zhong)。
對于異(yi)特(te)(te)(te)龍(long)(long)頭骨(gu)(gu)的(de)(de)(de)(de)有(you)限(xian)元分析顯示,異(yi)特(te)(te)(te)龍(long)(long)在攻(gong)擊獵物時,頭骨(gu)(gu)的(de)(de)(de)(de)上頜(he)骨(gu)(gu)-淚骨(gu)(gu)關節處(chu)會承(cheng)受(shou)非(fei)(fei)常大的(de)(de)(de)(de)反作用(yong)(yong)力。另外一點區別(bie)是,北美(mei)洲的(de)(de)(de)(de)暴(bao)龍(long)(long)類(lei)下(xia)頜(he)的(de)(de)(de)(de)齒(chi)骨(gu)(gu)和(he)(he)后面(mian)(mian)的(de)(de)(de)(de)骨(gu)(gu)塊(kuai)具(ju)有(you)較為(wei)靈(ling)活的(de)(de)(de)(de)關節,使得它(ta)們(men)的(de)(de)(de)(de)下(xia)頜(he)可以一定程度的(de)(de)(de)(de)內外擴展和(he)(he)收縮,而特(te)(te)(te)暴(bao)龍(long)(long)沒有(you)這樣(yang)(yang)的(de)(de)(de)(de)關節,使得它(ta)們(men)的(de)(de)(de)(de)下(xia)頜(he)更(geng)為(wei)一體化(hua)。造成這些區別(bie)的(de)(de)(de)(de)原(yuan)因(yin)主要(yao)是亞洲暴(bao)龍(long)(long)類(lei)和(he)(he)北美(mei)洲的(de)(de)(de)(de)暴(bao)龍(long)(long)類(lei)所面(mian)(mian)對的(de)(de)(de)(de)獵物不太一樣(yang)(yang)。晚(wan)白堊世北美(mei)洲的(de)(de)(de)(de)暴(bao)龍(long)(long)類(lei)主要(yao)捕食角龍(long)(long)類(lei)和(he)(he)鴨嘴龍(long)(long)類(lei),而亞洲的(de)(de)(de)(de)暴(bao)龍(long)(long)類(lei)則有(you)更(geng)多的(de)(de)(de)(de)機會捕獵大型蜥腳類(lei)恐龍(long)(long),比如泰(tai)坦巨龍(long)(long)類(lei)。目前(qian)還未有(you)研究使用(yong)(yong)模型化(hua)的(de)(de)(de)(de)方法(fa)精(jing)確計(ji)算過特(te)(te)(te)暴(bao)龍(long)(long)的(de)(de)(de)(de)咬合(he)力,但(dan)定性的(de)(de)(de)(de)比較認為(wei),特(te)(te)(te)暴(bao)龍(long)(long)與(yu)其(qi)它(ta)大型暴(bao)龍(long)(long)類(lei)一樣(yang)(yang)擁有(you)非(fei)(fei)常巨大的(de)(de)(de)(de)咬合(he)力。
菲利普.柯瑞等人在2003年(nian)的(de)(de)(de)(de)一項研(yan)究(jiu)中(zhong)報道了(le)2個(ge)可能屬于(yu)特暴龍(long)的(de)(de)(de)(de)腳(jiao)印(yin)。這些腳(jiao)印(yin)非常巨大,有(you)61厘米長,保存了(le)第三趾爪(zhua)在地(di)上(shang)留(liu)(liu)下的(de)(de)(de)(de)痕跡。同時放大觀察腳(jiao)印(yin)還可以看到里面的(de)(de)(de)(de)皮(pi)膚印(yin)痕。皮(pi)膚印(yin)痕中(zhong)的(de)(de)(de)(de)每個(ge)鱗(lin)片大約有(you)2毫(hao)米寬。另外一件(jian)特暴龍(long)的(de)(de)(de)(de)骨架上(shang)也發現了(le)皮(pi)膚印(yin)痕。皮(pi)膚印(yin)痕保留(liu)(liu)在接近咽(yan)喉的(de)(de)(de)(de)位置,鱗(lin)片的(de)(de)(de)(de)寬度(du)約為(wei)2.4毫(hao)米。遺(yi)憾(han)的(de)(de)(de)(de)是這件(jian)標本已經(jing)被破(po)壞,無法對(dui)其進行更深入的(de)(de)(de)(de)研(yan)究(jiu)。
一(yi)件(jian)特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)標本(ben)保存(cun)了完整的(de)(de)(de)(de)顱(lu)腔。通過制作顱(lu)內模可以了解特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)腦結(jie)構。特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)腦結(jie)構與霸王龍(long)(long)(long)(long)(long)十分相似,其主(zhu)要的(de)(de)(de)(de)區別(bie)僅存(cun)在于個別(bie)腦神(shen)經基部的(de)(de)(de)(de)位(wei)置(zhi),如(ru)三叉神(shen)經和(he)(he)副神(shen)經。一(yi)只12米長的(de)(de)(de)(de)特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)腦體積約(yue)為(wei)184立方厘米。特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)和(he)(he)霸王龍(long)(long)(long)(long)(long)一(yi)樣擁(yong)有(you)非(fei)常(chang)大的(de)(de)(de)(de)嗅葉,這(zhe)(zhe)說(shuo)明它(ta)(ta)們(men)有(you)良好(hao)的(de)(de)(de)(de)嗅覺(jue)。同時它(ta)(ta)們(men)也(ye)(ye)(ye)具有(you)發達(da)(da)的(de)(de)(de)(de)犁(li)鼻器用(yong)于探測荷(he)爾蒙,這(zhe)(zhe)說(shuo)明特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)可能擁(yong)有(you)復雜的(de)(de)(de)(de)交(jiao)配行為(wei)。特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)聽神(shen)經也(ye)(ye)(ye)很發達(da)(da),說(shuo)明它(ta)(ta)們(men)有(you)較(jiao)好(hao)的(de)(de)(de)(de)聽力。特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)中腦頂(ding)部(midbrain tectum)不甚發達(da)(da),它(ta)(ta)的(de)(de)(de)(de)動(dong)眼(yan)神(shen)經和(he)(he)視(shi)神(shen)經也(ye)(ye)(ye)是如(ru)此。在頭骨的(de)(de)(de)(de)宏觀形態形態上,特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)的(de)(de)(de)(de)頭骨后端向外側擴展的(de)(de)(de)(de)較(jiao)少(shao),其雙(shuang)眼(yan)視(shi)覺(jue)疊加范圍不如(ru)霸王龍(long)(long)(long)(long)(long)。因(yin)此特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)(long)(long)在生(sheng)活(huo)的(de)(de)(de)(de)時候對嗅覺(jue)和(he)(he)聽覺(jue)的(de)(de)(de)(de)依賴要遠大于對視(shi)覺(jue)的(de)(de)(de)(de)依賴。
同北(bei)美的(de)(de)(de)大型暴(bao)(bao)(bao)龍(long)(long)(long)類一(yi)(yi)(yi)樣(yang),特暴(bao)(bao)(bao)龍(long)(long)(long)在(zai)生長發(fa)(fa)(fa)育過(guo)程中也(ye)伴隨著一(yi)(yi)(yi)系列的(de)(de)(de)形態變(bian)化(hua)。目前發(fa)(fa)(fa)現(xian)(xian)的(de)(de)(de)大多(duo)數(shu)特暴(bao)(bao)(bao)龍(long)(long)(long)標(biao)(biao)本(ben)屬于(yu)亞成年個體或(huo)成年個體,只有(you)很少的(de)(de)(de)幼年個體。2011年發(fa)(fa)(fa)表的(de)(de)(de)一(yi)(yi)(yi)件特暴(bao)(bao)(bao)龍(long)(long)(long)幼體標(biao)(biao)本(ben)使得古生物學(xue)家對于(yu)特暴(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)幼年時(shi)期有(you)了(le)更(geng)多(duo)了(le)解。這(zhe)(zhe)只幼體在(zai)死亡的(de)(de)(de)時(shi)候大約(yue)只有(you)2-3歲。與(yu)(yu)成年個體相(xiang)比,幼體的(de)(de)(de)頭骨較(jiao)為(wei)脆弱,牙齒也(ye)不(bu)夠強壯,說(shuo)明幼體的(de)(de)(de)獵物與(yu)(yu)成體有(you)較(jiao)大的(de)(de)(de)不(bu)同。與(yu)(yu)霸王龍(long)(long)(long)和(he)懼龍(long)(long)(long)不(bu)同的(de)(de)(de)是,這(zhe)(zhe)件幼年特暴(bao)(bao)(bao)龍(long)(long)(long)標(biao)(biao)本(ben)的(de)(de)(de)齒骨齒和(he)上(shang)頜(he)骨齒的(de)(de)(de)數(shu)量(liang)與(yu)(yu)成年個體的(de)(de)(de)數(shu)量(liang)一(yi)(yi)(yi)致(zhi),說(shuo)明特暴(bao)(bao)(bao)龍(long)(long)(long)在(zai)個體發(fa)(fa)(fa)育的(de)(de)(de)過(guo)程中不(bu)存在(zai)牙齒數(shu)量(liang)變(bian)化(hua)的(de)(de)(de)現(xian)(xian)象(xiang)。這(zhe)(zhe)一(yi)(yi)(yi)區別是否(fou)廣泛存在(zai)與(yu)(yu)亞洲和(he)北(bei)美洲的(de)(de)(de)其它(ta)暴(bao)(bao)(bao)龍(long)(long)(long)亞科(ke)物種還需(xu)(xu)要進一(yi)(yi)(yi)步檢驗。對于(yu)幼體鞏(gong)膜環(huan)的(de)(de)(de)研究表明,幼體可能(neng)具(ju)有(you)夜(ye)行的(de)(de)(de)習(xi)性(xing)。成年特暴(bao)(bao)(bao)龍(long)(long)(long)是否(fou)具(ju)有(you)夜(ye)行習(xi)性(xing)還需(xu)(xu)要更(geng)多(duo)的(de)(de)(de)化(hua)石證據支持。
特暴(bao)龍(long)(long)(long)(long)(long)(long)(long)是其所處生態(tai)系(xi)統中最大的(de)捕食者(zhe),與其共存的(de)植(zhi)食性(xing)恐(kong)(kong)龍(long)(long)(long)(long)(long)(long)(long)包(bao)括(kuo)蜥腳類的(de)耐(nai)梅(mei)蓋特龍(long)(long)(long)(long)(long)(long)(long)和(he)(he)后凹尾龍(long)(long)(long)(long)(long)(long)(long),甲(jia)龍(long)(long)(long)(long)(long)(long)(long)類的(de)美(mei)甲(jia)龍(long)(long)(long)(long)(long)(long)(long),腫頭龍(long)(long)(long)(long)(long)(long)(long)類的(de)傾(qing)頭龍(long)(long)(long)(long)(long)(long)(long),鴨嘴龍(long)(long)(long)(long)(long)(long)(long)類的(de)櫛龍(long)(long)(long)(long)(long)(long)(long)和(he)(he)巴思(si)缽氏龍(long)(long)(long)(long)(long)(long)(long),屬(shu)(shu)(shu)于鐮(lian)(lian)刀龍(long)(long)(long)(long)(long)(long)(long)類的(de)鐮(lian)(lian)刀龍(long)(long)(long)(long)(long)(long)(long),屬(shu)(shu)(shu)于似鳥龍(long)(long)(long)(long)(long)(long)(long)類似鵝龍(long)(long)(long)(long)(long)(long)(long),似雞龍(long)(long)(long)(long)(long)(long)(long)和(he)(he)體型巨大的(de)恐(kong)(kong)手(shou)龍(long)(long)(long)(long)(long)(long)(long)。肉食性(xing)恐(kong)(kong)龍(long)(long)(long)(long)(long)(long)(long)則包(bao)括(kuo)同屬(shu)(shu)(shu)于暴(bao)龍(long)(long)(long)(long)(long)(long)(long)亞科的(de)分支龍(long)(long)(long)(long)(long)(long)(long),屬(shu)(shu)(shu)于傷(shang)齒(chi)龍(long)(long)(long)(long)(long)(long)(long)類的(de)無聊龍(long)(long)(long)(long)(long)(long)(long),鴕鳥龍(long)(long)(long)(long)(long)(long)(long)和(he)(he)扎(zha)納巴扎(zha)爾龍(long)(long)(long)(long)(long)(long)(long),屬(shu)(shu)(shu)于竊(qie)蛋龍(long)(long)(long)(long)(long)(long)(long)類的(de)單(dan)足龍(long)(long)(long)(long)(long)(long)(long),耐(nai)梅(mei)蓋特母龍(long)(long)(long)(long)(long)(long)(long)和(he)(he)瑞欽龍(long)(long)(long)(long)(long)(long)(long)。
特(te)(te)暴(bao)龍曾登場于BBC的紀錄片(pian)《恐龍兇(xiong)面目(mu)》以及《與龍同行特(te)(te)輯:尋爪記》。近期它又在紀錄片(pian)《史前星球》中登場。
在電影方面,特暴(bao)龍(long)登場于韓(han)國的動(dong)畫電影《韓(han)半島(dao)的恐(kong)龍(long)》以及國產恐(kong)龍(long)動(dong)畫電影《恐(kong)龍(long)王》。
特暴龍位于食物鏈的頂端,是(shi)(shi)一(yi)種(zhong)頂級掠食動(dong)物。特暴龍在暴龍科中(zhong)的分類(lei)位置仍未(wei)確(que)定。以前(qian),有(you)(you)些(xie)科學家(jia)(jia)認為勇(yong)士特暴龍其實是(shi)(shi)北美洲暴龍的亞洲種(zhong)。如果(guo)屬實,將使(shi)特暴龍成為無效的分類(lei)。即使(shi)特暴龍與暴龍不是(shi)(shi)同種(zhong)動(dong)物,它們被認為有(you)(you)接近的親緣關系。有(you)(you)些(xie)科學家(jia)(jia)認為,同樣發(fa)現(xian)于蒙古的分支龍,是(shi)(shi)特暴龍的近親。
特暴龍的(de)(de)化(hua)石記錄保存良好,已有(you)數十(shi)個標本,包含至(zhi)少5個完整(zheng)的(de)(de)頭(tou)顱骨與骨骸。這些化(hua)石讓科學(xue)家得以研(yan)究它們的(de)(de)種系發生學(xue)、頭(tou)部(bu)力學(xue)、以及(ji)腦(nao)部(bu)結(jie)構。
特暴龍(long)屬于暴龍(long)科(ke)(ke)的(de)(de)暴龍(long)亞科(ke)(ke)。該亞科(ke)(ke)還(huan)包(bao)含較(jiao)(jiao)早期的(de)(de)懼龍(long)、較(jiao)(jiao)晚期的(de)(de)暴龍(long),都(dou)發現于北美洲(zhou),可能還(huan)有蒙古的(de)(de)分支龍(long)。暴龍(long)亞科(ke)(ke)包(bao)含親緣關系較(jiao)(jiao)接近(jin)暴龍(long),而(er)離艾伯塔(ta)龍(long)較(jiao)(jiao)遠的(de)(de)物(wu)種;與艾伯塔(ta)龍(long)亞科(ke)(ke)相(xiang)比(bi),暴龍(long)亞科(ke)(ke)的(de)(de)體格較(jiao)(jiao)重型(xing),頭(tou)顱骨的(de)(de)比(bi)例較(jiao)(jiao)大,以及較(jiao)(jiao)長的(de)(de)股骨。
勇士特暴龍最初被視為暴龍的一個種,某些近年的分類也支持這個說(shuo)法。其他(ta)的(de)科學(xue)家則將它們(men)列為(wei)獨立的(de)屬,并(bing)為(wei)暴(bao)(bao)(bao)龍(long)(long)(long)的(de)姐妹(mei)分類(lei)單(dan)元(yuan)。在(zai)2003年(nian),一(yi)個(ge)(ge)親緣分支分類(lei)法研究提出(chu)(chu)分支龍(long)(long)(long)是(shi)特(te)暴(bao)(bao)(bao)龍(long)(long)(long)的(de)近親,因(yin)為(wei)它們(men)具有(you)其他(ta)暴(bao)(bao)(bao)龍(long)(long)(long)亞科沒有(you)的(de)頭(tou)部(bu)特(te)征。如果屬實(shi),將排(pai)除特(te)暴(bao)(bao)(bao)龍(long)(long)(long)是(shi)暴(bao)(bao)(bao)龍(long)(long)(long)的(de)一(yi)個(ge)(ge)異名的(de)可能性,并(bing)顯示暴(bao)(bao)(bao)龍(long)(long)(long)亞科在(zai)北美洲與亞洲演化(hua)出(chu)(chu)個(ge)(ge)別的(de)支系。分支龍(long)(long)(long)的(de)唯(wei)一(yi)標本具有(you)幼年(nian)體(ti)的(de)特(te)征,但牙(ya)齒(chi)數量較(jiao)多(duo),約76到(dao)78顆,而且口鼻部(bu)上面有(you)獨特(te)的(de)低矮骨質瘤,因(yin)此并(bing)非特(te)暴(bao)(bao)(bao)龍(long)(long)(long)的(de)幼年(nian)體(ti)。
特(te)暴(bao)龍(long)過去生存(cun)于潮濕(shi)的泛(fan)濫平(ping)原,布滿者河道。在(zai)中國(guo)的分布包(bao)括黑龍(long)江,河南,山東,廣東,云(yun)南,內蒙(meng)古等地,國(guo)外主(zhu)要是(shi)蒙(meng)古。在(zai)中國(guo)黑龍(long)江,河南,山東,廣東的所謂中國(guo)"暴(bao)龍(long)"其實都是(shi)特(te)暴(bao)龍(long)。
特(te)暴(bao)(bao)(bao)(bao)龍(long)是(shi)最(zui)大型(xing)(xing)(xing)的(de)(de)(de)(de)暴(bao)(bao)(bao)(bao)龍(long)科動(dong)物之一(yi)(yi),但略小于(yu)暴(bao)(bao)(bao)(bao)龍(long)。已(yi)知最(zui)大型(xing)(xing)(xing)的(de)(de)(de)(de)個體(ti)身長(chang)12米,頭部(bu)(bu)離(li)地面(mian)約4.2米。一(yi)(yi)般體(ti)重3噸(dun)到5噸(dun),最(zui)大的(de)(de)(de)(de)化(hua)石(shi)體(ti)重可達到7.5噸(dun)。如同大部(bu)(bu)分已(yi)知的(de)(de)(de)(de)暴(bao)(bao)(bao)(bao)龍(long)科恐龍(long),特(te)暴(bao)(bao)(bao)(bao)龍(long)是(shi)種大型(xing)(xing)(xing)、二(er)足(zu)掠食動(dong)物,重達數(shu)噸(dun),擁(yong)(yong)有數(shu)十顆(ke)大型(xing)(xing)(xing)、銳(rui)利的(de)(de)(de)(de)牙(ya)齒。特(te)暴(bao)(bao)(bao)(bao)龍(long)的(de)(de)(de)(de)下頜有特(te)殊的(de)(de)(de)(de)接合構(gou)造。另外,就(jiu)前(qian)肢/身體(ti)比例而言,特(te)暴(bao)(bao)(bao)(bao)龍(long)擁(yong)(yong)有暴(bao)(bao)(bao)(bao)龍(long)科中最(zui)小型(xing)(xing)(xing)的(de)(de)(de)(de)前(qian)肢。特(te)暴(bao)(bao)(bao)(bao)龍(long)是(shi)種大型(xing)(xing)(xing)、二(er)足(zu)掠食動(dong)物,重達數(shu)噸(dun)。特(te)暴(bao)(bao)(bao)(bao)龍(long)唯一(yi)(yi)一(yi)(yi)個有效(xiao)種的(de)(de)(de)(de)體(ti)型(xing)(xing)(xing):勇士特(te)暴(bao)(bao)(bao)(bao)龍(long)Tarbosaurus bataar 8-12米。
暴(bao)龍科的身體外形(xing)差(cha)異不大。特暴(bao)龍的頸(jing)部為S狀彎曲,其(qi)余的脊(ji)柱,包含尾巴,與(yu)地面保持者水平的姿(zi)態。
就前肢(zhi)/身體(ti)(ti)比例而言,特暴龍擁有暴龍科中最小型的(de)(de)(de)前肢(zhi)。有兩根(gen)迷(mi)你的(de)(de)(de)手(shou)指。后肢(zhi)長(chang)而粗(cu)厚(hou),將身體(ti)(ti)支撐為二足的(de)(de)(de)步態,上(shang)有三根(gen)腳趾。長(chang)而重(zhong)的(de)(de)(de)尾巴可以(yi)平衡頭部(bu)與胸部(bu)的(de)(de)(de)重(zhong)量,將重(zhong)心保持(chi)在臀部(bu)。
在2003年,特暴(bao)(bao)龍(long)的(de)顱骨(gu)首次(ci)經過完(wan)整的(de)研究。科學(xue)家們發現(xian)特暴(bao)(bao)龍(long)與北(bei)美洲暴(bao)(bao)龍(long)科之間(jian)有(you)幾個顯著的(de)差(cha)異。這些差(cha)異與顱骨(gu)在咬(yao)合(he)時,力(li)量(liang)的(de)傳遞(di)有(you)關。當特暴(bao)(bao)龍(long)的(de)上(shang)(shang)(shang)頜(he)咬(yao)住物(wu)體(ti)時,力(li)量(liang)從(cong)上(shang)(shang)(shang)頜(he)骨(gu)傳遞(di)到(dao)上(shang)(shang)(shang)頜(he)周遭的(de)顱骨(gu)。而北(bei)美洲暴(bao)(bao)龍(long)科咬(yao)合(he)時,力(li)量(liang)從(cong)上(shang)(shang)(shang)頜(he)骨(gu)傳遞(di)到(dao)口鼻(bi)部上(shang)(shang)(shang)方的(de)固定鼻(bi)骨(gu),然后再經由鼻(bi)骨(gu)與淚骨(gu)間(jian)的(de)骨(gu)質連結,傳遞(di)到(dao)淚骨(gu)。
特暴龍的(de)(de)鼻骨(gu)(gu)(gu)與淚(lei)(lei)骨(gu)(gu)(gu)間缺乏骨(gu)(gu)(gu)質(zhi)的(de)(de)連(lian)結。但上(shang)頜(he)骨(gu)(gu)(gu)的(de)(de)后方有個大型突起,楔合入淚(lei)(lei)骨(gu)(gu)(gu)內;而北(bei)美洲暴龍的(de)(de)上(shang)頜(he)骨(gu)(gu)(gu)后突很小。這個特征顯示,咬合的(de)(de)力量從特暴龍的(de)(de)上(shang)頜(he)骨(gu)(gu)(gu)直接(jie)傳遞到淚(lei)(lei)骨(gu)(gu)(gu)。而淚(lei)(lei)骨(gu)(gu)(gu)與額(e)骨(gu)(gu)(gu)、前(qian)額(e)骨(gu)(gu)(gu)之間更為牢(lao)固(gu)(gu)。由于上(shang)頜(he)骨(gu)(gu)(gu)、淚(lei)(lei)骨(gu)(gu)(gu)、額(e)骨(gu)(gu)(gu)、前(qian)額(e)骨(gu)(gu)(gu)之間牢(lao)牢(lao)地(di)固(gu)(gu)定者,使得上(shang)頜(he)非常堅固(gu)(gu)。
另一個(ge)主要的(de)(de)差別則(ze)是(shi)堅(jian)固的(de)(de)下頜(he)。許多獸腳(jiao)類(lei)恐(kong)龍,包括北美洲暴龍科在內,下頜(he)的(de)(de)齒骨(gu)與后面骨(gu)頭(tou)間有(you)靈活(huo)的(de)(de)關節(jie)。特暴龍的(de)(de)隅骨(gu)側邊(bian)棱脊連(lian)接者齒骨(gu)后方(fang)的(de)(de)方(fang)形突(tu),使(shi)它(ta)們(men)的(de)(de)下頜(he)無法靈活(huo)外內扳動。
有些科(ke)學(xue)家(jia)提出(chu)假設,認為特暴龍(long)的(de)(de)(de)堅硬頭部(bu)是種適(shi)應演(yan)化,用(yong)來(lai)獵殺耐梅(mei)蓋(gai)特組的(de)(de)(de)大(da)型蜥腳(jiao)類(lei)恐龍(long),泰坦巨(ju)龍(long)類(lei),因為晚白堊紀的(de)(de)(de)北美洲并沒有如此巨(ju)大(da)的(de)(de)(de)恐龍(long)存在。這種頭部(bu)力(li)學(xue)的(de)(de)(de)差異也影響(xiang)了暴龍(long)科(ke)的(de)(de)(de)種系發生學(xue)。同樣發現于(yu)蒙(meng)古的(de)(de)(de)分支(zhi)龍(long),也具(ju)有類(lei)似特暴龍(long)的(de)(de)(de)頭部(bu)力(li)學(xue)特征,這顯示(shi)暴龍(long)并非特暴龍(long)的(de)(de)(de)近親。特暴龍(long)與(yu)暴龍(long)的(de)(de)(de)相似處可能導因于(yu)它們巨(ju)大(da)的(de)(de)(de)體型,是平(ping)行演(yan)化的(de)(de)(de)結果。
一個在1948年(nian)發(fa)現的(de)(de)顱骨,一度被歸類于(yu)蛇(she)發(fa)女怪龍的(de)(de)G. lancinator,因為具有顱腔,有助于(yu)科學家(jia)了(le)(le)(le)解特暴龍的(de)(de)腦(nao)部(bu)結(jie)(jie)構(gou)。在1965年(nian),馬列夫(fu)制作了(le)(le)(le)一個石膏顱腔模型(xing)(xing),并(bing)做了(le)(le)(le)初步的(de)(de)腦(nao)部(bu)形狀(zhuang)檢查。在2005年(nian),謝(xie)爾蓋·薩伐黎耶夫(fu)(Sergei V. Saveliev)制作了(le)(le)(le)一個聚氨酯顱腔模型(xing)(xing),并(bing)做了(le)(le)(le)更詳細的(de)(de)腦(nao)部(bu)結(jie)(jie)構(gou)與功能研究(jiu)。
根據暴(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)腦(nao)部(bu)(bu)結構研(yan)究,它們(men)(men)兩(liang)者的(de)(de)(de)(de)腦(nao)部(bu)(bu)結構相似(si),只有某些腦(nao)神(shen)(shen)經(jing)(jing)根部(bu)(bu)的(de)(de)(de)(de)位置(zhi)不一樣,包含三叉神(shen)(shen)經(jing)(jing)與(yu)(yu)(yu)副神(shen)(shen)經(jing)(jing)。暴(bao)(bao)(bao)龍(long)(long)(long)科的(de)(de)(de)(de)腦(nao)部(bu)(bu)較(jiao)類(lei)似(si)鱷魚與(yu)(yu)(yu)其(qi)他爬(pa)蟲類(lei),而(er)(er)較(jiao)不類(lei)似(si)鳥(niao)類(lei)。特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)腦(nao)部(bu)(bu)體(ti)(ti)積為184立(li)方厘米。腦(nao)部(bu)(bu)的(de)(de)(de)(de)大(da)型嗅球(qiu)、末端(duan)神(shen)(shen)經(jing)(jing)、嗅神(shen)(shen)經(jing)(jing),顯(xian)(xian)示特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)具(ju)有靈敏的(de)(de)(de)(de)嗅覺(jue),這(zhe)點如同(tong)暴(bao)(bao)(bao)龍(long)(long)(long)。犁鼻(bi)球(qiu)大(da)型,而(er)(er)且與(yu)(yu)(yu)嗅球(qiu)分離,顯(xian)(xian)示可以(yi)感應費洛蒙的(de)(de)(de)(de)犁鼻(bi)器發(fa)展(zhan)良(liang)好(hao)。這(zhe)個特(te)(te)征顯(xian)(xian)示特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)可能(neng)具(ju)有復雜的(de)(de)(de)(de)求偶行為。聽神(shen)(shen)經(jing)(jing)也很大(da),顯(xian)(xian)示它們(men)(men)的(de)(de)(de)(de)聽力(li)(li)很好(hao),可能(neng)用在(zai)聲音的(de)(de)(de)(de)溝通(tong)與(yu)(yu)(yu)警告(gao)上。聽神(shen)(shen)經(jing)(jing)連接者發(fa)展(zhan)良(liang)好(hao)的(de)(de)(de)(de)前庭系統,顯(xian)(xian)示它們(men)(men)的(de)(de)(de)(de)平衡感與(yu)(yu)(yu)協(xie)調性很好(hao)。相反地,與(yu)(yu)(yu)視力(li)(li)有關的(de)(de)(de)(de)腦(nao)部(bu)(bu)區域與(yu)(yu)(yu)神(shen)(shen)經(jing)(jing)較(jiao)小。爬(pa)蟲類(lei)的(de)(de)(de)(de)中(zhong)腦(nao)頂蓋連接控制眼球(qiu)的(de)(de)(de)(de)視神(shen)(shen)經(jing)(jing)與(yu)(yu)(yu)動眼神(shen)(shen)經(jing)(jing),是用來處(chu)理(li)視力(li)(li)訊息(xi),但特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)中(zhong)腦(nao)頂蓋非常小。暴(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)眼睛朝(chao)前,因(yin)此具(ju)有一定程(cheng)度的(de)(de)(de)(de)立(li)體(ti)(ti)視覺(jue);但特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)的(de)(de)(de)(de)顱骨狹窄,眼睛朝(chao)向兩(liang)側(ce),如同(tong)典型的(de)(de)(de)(de)暴(bao)(bao)(bao)龍(long)(long)(long)科動物。這(zhe)些特(te)(te)征顯(xian)(xian)示特(te)(te)暴(bao)(bao)(bao)龍(long)(long)(long)較(jiao)依(yi)靠(kao)嗅覺(jue)與(yu)(yu)(yu)聽覺(jue),而(er)(er)非視覺(jue)。
已知最(zui)大(da)型的(de)特暴(bao)龍(long)顱(lu)骨長(chang)度超過1.3米,小于(yu)暴(bao)龍(long)和(he)諸城暴(bao)龍(long)以及懼龍(long),但大(da)于(yu)其它的(de)暴(bao)龍(long)科(ke)。如同暴(bao)龍(long),特暴(bao)龍(long)的(de)顱(lu)骨高大(da),前段(duan)狹窄。顱(lu)骨后段(duan)擴(kuo)張幅度不大(da),意味著特暴(bao)龍(long)所(suo)以不如暴(bao)龍(long)擁(yong)有的(de)立(li)體(ti)視覺好。
顱(lu)骨的(de)大(da)型洞孔可(ke)減(jian)輕重量。特(te)暴龍(long)的(de)下頜有(you)特(te)殊的(de)接合構造。
顎(e)部(bu)有60到64顆牙(ya)齒,略少于暴(bao)(bao)龍(long),但(dan)大于其(qi)它(ta)(ta)體型較小的(de)(de)暴(bao)(bao)龍(long)科(ke)(ke),例如蛇發女怪龍(long)與分支龍(long)。大部(bu)分的(de)(de)牙(ya)齒橫(heng)(heng)剖(pou)面為橢(tuo)圓形(xing),而前上顎(e)骨的(de)(de)牙(ya)齒橫(heng)(heng)剖(pou)面為D形(xing)。暴(bao)(bao)龍(long)科(ke)(ke)都具有這種異型齒特(te)征。上顎(e)骨的(de)(de)牙(ya)齒最長(chang)(chang),齒冠長(chang)(chang)達85公厘。如同(tong)其(qi)它(ta)(ta)大型暴(bao)(bao)龍(long)科(ke)(ke)與現(xian)代(dai)的(de)(de)科(ke)(ke)莫多龍(long),特(te)暴(bao)(bao)龍(long)的(de)(de)幼年(nian)與近(jin)成年(nian)個(ge)體的(de)(de)身上發現(xian)過有齒痕,符(fu)合(he)成年(nian)個(ge)體或其(qi)它(ta)(ta)較小型獸腳(jiao)類恐龍(long)的(de)(de)牙(ya)齒。
特(te)暴龍(long)與分支龍(long)的下(xia)顎外(wai)側各有一道棱脊,從隅骨(gu)延(yan)伸(shen)到(dao)齒骨(gu)后方,形成相扣的結構。其它暴龍(long)科動物缺乏這道棱脊,因(yin)此(ci)下(xia)顎更為(wei)靈活。
特暴(bao)龍位(wei)于食物鏈的(de)(de)頂端,是一(yi)種頂級掠(lve)食者(zhe),可能以大型恐龍為食,例如鴨嘴龍類的(de)(de)櫛(zhi)龍,或(huo)是蜥腳類的(de)(de)納摩蓋吐龍。
成年(nian)特暴龍(long)(long)(long)(long)可能與其它小型獸(shou)腳(jiao)類(lei)(lei)恐(kong)(kong)(kong)龍(long)(long)(long)(long)有少許(xu)競爭,例如(ru)傷齒(chi)龍(long)(long)(long)(long)科(ke)的(de)(de)(de)(de)無聊(liao)龍(long)(long)(long)(long)、鴕(tuo)鳥(niao)龍(long)(long)(long)(long)、蜥鳥(niao)龍(long)(long)(long)(long),以及偷蛋(dan)龍(long)(long)(long)(long)下目的(de)(de)(de)(de)單足龍(long)(long)(long)(long)、瑞欽龍(long)(long)(long)(long),或(huo)者還有小掠龍(long)(long)(long)(long),一(yi)種有時被認為是基底暴龍(long)(long)(long)(long)超科(ke)的(de)(de)(de)(de)恐(kong)(kong)(kong)龍(long)(long)(long)(long)。其它的(de)(de)(de)(de)獸(shou)腳(jiao)類(lei)(lei)恐(kong)(kong)(kong)龍(long)(long)(long)(long),包含:巨大的(de)(de)(de)(de)鐮刀龍(long)(long)(long)(long)、似鳥(niao)龍(long)(long)(long)(long)下目的(de)(de)(de)(de)似鵝(e)龍(long)(long)(long)(long)、似雞龍(long)(long)(long)(long)、恐(kong)(kong)(kong)手龍(long)(long)(long)(long),鐮刀龍(long)(long)(long)(long)可能是草食(shi)(shi)性,而(er)上述似鳥(niao)龍(long)(long)(long)(long)類(lei)(lei)恐(kong)(kong)(kong)龍(long)(long)(long)(long)可能為雜食(shi)(shi)性,以小型動物為食(shi)(shi),不(bu)會跟特暴龍(long)(long)(long)(long)競爭食(shi)(shi)物。
在2001年,布(bu)魯斯·羅斯柴爾(er)德(Bruce Rothschild)等人發表一份獸(shou)腳類恐龍的(de)壓(ya)(ya)力(li)性(xing)(xing)骨(gu)(gu)(gu)折(zhe)與肌(ji)腱撕裂傷研(yan)究(jiu),并研(yan)究(jiu)它們的(de)行為模(mo)式。壓(ya)(ya)力(li)性(xing)(xing)骨(gu)(gu)(gu)折(zhe)較(jiao)常(chang)導(dao)因(yin)于(yu)(yu)習(xi)慣性(xing)(xing)動作,較(jiao)少來自于(yu)(yu)外力(li)沖擊。他們研(yan)究(jiu)18個特暴龍的(de)腳掌骨(gu)(gu)(gu)骼,沒有(you)(you)發現(xian)壓(ya)(ya)力(li)性(xing)(xing)骨(gu)(gu)(gu)折(zhe)的(de)跡象(xiang)(xiang);而(er)在10個被(bei)研(yan)究(jiu)的(de)手(shou)掌骨(gu)(gu)(gu)頭(tou)(tou)中,其中一個手(shou)部骨(gu)(gu)(gu)頭(tou)(tou)有(you)(you)發現(xian)壓(ya)(ya)力(li)性(xing)(xing)骨(gu)(gu)(gu)折(zhe)的(de)跡象(xiang)(xiang)。腳部骨(gu)(gu)(gu)頭(tou)(tou)的(de)壓(ya)(ya)力(li)性(xing)(xing)骨(gu)(gu)(gu)折(zhe),可以歸咎于(yu)(yu)奔跑(pao)、長途遷(qian)徙。手(shou)部骨(gu)(gu)(gu)頭(tou)(tou)的(de)壓(ya)(ya)力(li)性(xing)(xing)骨(gu)(gu)(gu)折(zhe),極(ji)可能導(dao)因(yin)于(yu)(yu)捕抓(zhua)獵(lie)物而(er)受傷。這些壓(ya)(ya)力(li)性(xing)(xing)骨(gu)(gu)(gu)折(zhe)、肌(ji)腱撕裂傷跡象(xiang)(xiang),顯(xian)示特暴龍常(chang)因(yin)為捕抓(zhua)獵(lie)物而(er)受傷,而(er)非(fei)吞食尸體。
在(zai)1946年,一個蘇聯與(yu)蒙古(gu)挖掘團隊在(zai)蒙古(gu)南戈壁省(sheng)的(de)耐梅蓋特組發現一個接近(jin)完整的(de)絕大部分大型頭顱骨與(yu)一些脊(ji)椎(zhui)骨,屬于一個大型的(de)獸腳類恐龍。
在(zai)1955年,蘇聯古生物學家葉(xie)甫根尼·馬列夫(Evgeny Maleev)將這正模標本(PIN 551-1)建立為(wei)(wei)暴(bao)龍的一個種,勇士暴(bao)龍(Tyrannosaurus bataar)。種名(ming)баатар/baatar在(zai)蒙古語中意為(wei)(wei)"勇士",但(dan)被誤拼(pin)為(wei)(wei)bataar。
同年(nian)(nian),馬列夫將3個獸腳類的化(hua)石敘述(shu)并命(ming)名,三(san)者都(dou)是頭顱骨,外加部(bu)分身(shen)體,全是同一個挖掘團隊在1948到49年(nian)(nian)間(jian)發現的。第一個(編(bian)(bian)號PIN 551-2)被建立為新屬(shu),埃夫雷(lei)莫夫特暴(bao)龍(T. efremovi),ταρβο?/tarbos在希臘文文中意為"恐怖(bu)的"、"敬畏(wei)的",而σαυρο?/saurus意為"蜥蜴(yi)",種名是以蘇聯古(gu)生物學家兼科幻(huan)小說作(zuo)者伊凡(fan)·埃夫雷(lei)莫夫(Ivan Yefremov)為名。另(ling)外兩個(編(bian)(bian)號PIN 553-1與PIN 552-2)被歸類于北(bei)美洲的蛇發女怪龍,G. lancinator與G. novojilovi。這三(san)個標本(ben)都(dou)小于1946年(nian)(nian)發現的標本(ben)。
自(zi)從(cong)蘇聯與蒙古(gu)的(de)(de)挖(wa)掘(jue)團隊在40年代的(de)(de)挖(wa)掘(jue)過后,一個波蘭與蒙古(gu)的(de)(de)挖(wa)掘(jue)團隊再度(du)回到戈壁沙漠挖(wa)掘(jue),從(cong)1963年持續到1971年,發現了許(xu)多新(xin)的(de)(de)化石(shi),并在耐梅蓋(gai)特組(zu)發現了特暴龍的(de)(de)新(xin)標本。
在(zai)1965年(nian),蘇聯古生(sheng)物學(xue)家阿納托利·康斯坦丁諾(nuo)維(wei)奇·羅特杰(jie)斯特文(wen)斯基(Anatoly Konstantinovich Rozhdestvensky)認為馬列夫所發現(xian)的(de)標本,其實是同種動物的(de)不(bu)同生(sheng)長階(jie)段,而且不(bu)同于北美洲的(de)暴龍(long)。
1979年,董(dong)枝明(ming)在(zai)根據欒(luan)(luan)川(chuan)縣發掘(jue)到五顆(ke)大型牙齒(chi)命名為巒川(chuan)暴(bao)(bao)龍(long)(long)(Tyrannosaurus luanchuanensis)和霸王龍(long)(long)相似(si)種(zhong)(Tyrannosaurus cf. T. rex )的(de)(de)動(dong)物后來就(jiu)被(bei)改為欒(luan)(luan)川(chuan)特(te)暴(bao)(bao)龍(long)(long)等,而欒(luan)(luan)川(chuan)特(te)暴(bao)(bao)龍(long)(long)實際(ji)也(ye)不能成立,因(yin)為后來的(de)(de)研究(jiu)把它(ta)歸入了勇士(shi)特(te)暴(bao)(bao)龍(long)(long),認為它(ta)實際(ji)只是勇士(shi)特(te)暴(bao)(bao)龍(long)(long)的(de)(de)青年個體。在(zai)欒(luan)(luan)川(chuan)特(te)暴(bao)(bao)龍(long)(long)發現(xian)之前(qian),廣東河(he)源發現(xian)過特(te)暴(bao)(bao)龍(long)(long)未定(ding)種(zhong)的(de)(de)化石。發現(xian)欒(luan)(luan)川(chuan)特(te)暴(bao)(bao)龍(long)(long)的(de)(de)秋扒組(zu)巖石,秋扒組(zu)巖石是分(fen)布在(zai)潭頭盆地欒(luan)(luan)川(chuan)縣一帶的(de)(de)白堊紀(ji)晚(wan)期地層,是由(you)紫紅色(se)的(de)(de)砂泥巖構成,最底部含有磚紅色(se)的(de)(de)礫石,其他發現(xian)過特(te)暴(bao)(bao)龍(long)(long)的(de)(de)地層也(ye)都屬于(yu)白堊紀(ji)晚(wan)期。
在1986年,雖然(ran)已經發現(xian)許多特(te)(te)(te)(te)(te)(te)暴龍(long)標本(ben),但很少已確(que)定的(de)明確(que)資料,特(te)(te)(te)(te)(te)(te)暴龍(long)被(bei)推論(lun)與其他(ta)暴龍(long)科(ke)恐龍(long)有許多共同特(te)(te)(te)(te)(te)(te)征(zheng)。這些相(xiang)近的(de)特(te)(te)(te)(te)(te)(te)征(zheng)讓某(mou)些科(ke)學家提(ti)出當時的(de)北美洲與歐亞大陸之間有可能(neng)的(de)連結,也許是陸橋。他(ta)將四個標本(ben)與新(xin)發現(xian)的(de)化石(shi),統合為勇士(shi)特(te)(te)(te)(te)(te)(te)暴龍(long)(T.bataar)。后來的(de)研究人(ren)員(yuan)都同意羅特(te)(te)(te)(te)(te)(te)杰斯(si)特(te)(te)(te)(te)(te)(te)文(wen)斯(si)基(ji)的(de)分類,包(bao)含(han)馬(ma)列夫(fu)在內,但少數科(ke)學家使用埃夫(fu)雷(lei)莫夫(fu)特(te)(te)(te)(te)(te)(te)暴龍(long),而非勇士(shi)特(te)(te)(te)(te)(te)(te)暴龍(long)。
在1992年,美國古(gu)生物學(xue)家肯尼思·卡彭特(Kenneth Carpenter)重新檢(jian)驗這些(xie)化(hua)石,他根據頭(tou)顱骨的(de)相似(si)處,提(ti)出這些(xie)化(hua)石大部分屬于暴(bao)龍,并將(jiang)勇士(shi)特暴(bao)龍改回勇士(shi)暴(bao)龍;而蛇(she)發女怪龍的(de)G. novojilovi是個較(jiao)小型的(de)暴(bao)龍科動物,另外建(jian)立為(wei)馬(ma)列(lie)夫(fu)龍(Maleevosaurus novojilovi)。
在1995年,喬治·奧利舍夫(fu)斯基(George Olshevsky)建立(li)了(le)勇士成(cheng)吉思(si)汗(han)龍(long)(long)(Jenghizkhan bataar),取(qu)代勇士特暴龍(long)(long),并以成(cheng)吉思(si)汗(han)為名;他并提(ti)出埃夫(fu)雷(lei)莫夫(fu)特暴龍(long)(long)、馬列夫(fu)龍(long)(long)、勇士成(cheng)吉思(si)汗(han)龍(long)(long)是三個獨(du)立(li)的屬,都生存于相同時期的耐梅蓋(gai)特組。
在1999年(nian),一個研(yan)究提(ti)出馬列夫龍是特(te)暴(bao)(bao)龍的(de)幼年(nian)個體。自從(cong)1999年(nian)以來,所有的(de)研(yan)究認為(wei)這些化石皆為(wei)同種動物,勇士特(te)暴(bao)(bao)龍或是勇士暴(bao)(bao)龍。
在1993年(nian)到1998年(nian),日本與蒙古(gu)的(de)(de)挖掘團隊,以及21世紀(ji)初(chu),加拿大古(gu)生物學家菲力·柯爾(Phil Currie)的(de)(de)挖掘小組,也發現(xian)了(le)許多特暴龍(long)化石。不算上私人收藏的(de)(de)標本,已發現(xian)了(le)至少37個標本,包(bao)含超(chao)過15個完(wan)整和部分的(de)(de)頭顱(lu)骨(gu)化石,與至少5個完(wan)整的(de)(de)顱(lu)后(hou)骨(gu)骸。
21世紀初,廣東河源盆地也出(chu)土過大型(xing)獸腳類的大型(xing)暴龍類牙齒化(hua)石,推(tui)測可(ke)能是特暴龍。
特(te)暴(bao)(bao)(bao)龍的(de)大部分化石(shi),是成(cheng)年或(huo)亞成(cheng)年個(ge)(ge)體(ti)(ti),很少發(fa)現(xian)幼(you)年個(ge)(ge)體(ti)(ti)的(de)化石(shi)。在2006年發(fa)現(xian)的(de)一個(ge)(ge)幼(you)年個(ge)(ge)體(ti)(ti)身體(ti)(ti)骨骼,帶有完整的(de)頭顱骨,頭部長(chang)度為29厘米,使得(de)古生物學家可以(yi)研究特(te)暴(bao)(bao)(bao)龍的(de)生長(chang)模(mo)式。這(zhe)個(ge)(ge)幼(you)年個(ge)(ge)體(ti)(ti)化石(shi),死亡時的(de)年齡大約是2到3歲。與成(cheng)年個(ge)(ge)體(ti)(ti)相比,這(zhe)個(ge)(ge)幼(you)年頭顱骨的(de)結構虛(xu)弱,牙齒較細(xi),顯示特(te)暴(bao)(bao)(bao)龍的(de)幼(you)年個(ge)(ge)體(ti)(ti)、成(cheng)年個(ge)(ge)體(ti)(ti)占據不同的(de)生態位,以(yi)免(mian)競爭(zheng)相同食(shi)物來源。